Three unlikely friends, "Sid the Land Sloth" (voiced by John Leguizamo), "Manford 'Manny' the Wolly Mamoth" (voiced by Ray Ramono) and "Diego the Saber-tooth Tiger" (voiced by Dennis Leary) have discovered that the ice damn keeping the area they and other species live dry is melting. There claims are confirmed by a vulture.
The residents of this area go on a journey to a natural boat made out of a large tree for safety. Along the way, they meet "Ellie" (voiced by Queen Latifah), a female Wolly Mamoth who believes she is a possum, and her two "brothers", and actual possums, "Eddie" (voiced by Josh Peck) and "Crash" (voiced by Seann William Scott).
The group then go on an adventure to the boat, while trying to dodge predators that are following them.
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